(Lesquereux) Brotherus in H. G. A. Engler and K. Prantl, Nat. Pflanzenfam. 214 [I,3]: 431. 1902.
Lesquereux, Mem. Calif. Acad. Sci. 1: 12. 1868;
Cardot & Thériot
short-elliptic or long-obovate, sometimes ovate, apex very broadly acute to rounded, short- to long-awned, margins very broadly recurved or revolute to near apex, not bordered; costa usually long-excurrent, lacking an adaxial pad of cells, distally narrow, ca. 3-4 cells across the convex adaxial surface; distal laminal cells hexagonal, 10-15 µm wide, 1:1, densely papillose with 2-fid papillae.
apparently dioicous but more probably rhizautoicous, occasionally autoicous.
length ca. 0.7-1.4(-2) cm.
stegocarpic, not systylius, cylindric, erect and nearly straight, urn length ca. (1.7-)2.5-3.5 mm; peristome 800-1000 µm, teeth of 32 filaments twisted about one full turn, basal membrane 150-200 µm; operculum 1-1.5 mm.
ca. 10-13 µm, spheric, finely papillose.
Calcareous soil; low to moderate elevations; Ariz., Calif., Colo., Idaho, Nev., Oreg., Utah, Wash.; Mexico (Baja California, Sonora).
is much like
, but differs in the soil substrate, absence of a leaf border (common in
), the apparently dioicous inflorescence, the commonly very long (to 5 mm) operculate capsule, and the well-developed basal membrane of the peristome. According to B. D. Mishler (1994),
in Mexico intergrades with
in capsule length, height of the basal membrane, and development of a leaf border.