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Raja clavata Linnaeus, 1758
THORNBACK RAY
Life   Vertebrata   Fish   Rajidae   Raja

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

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Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata

Raja clavata
© Copyright Malcolm Storey 2011-2118 · 3
Raja clavata
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Common name (e.g. trout)

Genus + Species (e.g. Gadus morhua)

Raja clavata Linnaeus , 1758

Thornback ray Add your observation in Fish Watcher
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Raja clavata   AquaMaps   Data sources: GBIF OBIS
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Pictures | Videos |     Google image Image of Raja clavata (Thornback ray) Raja clavata
Picture by Stergiou, K.I.

Classification / Names Common names | Synonyms | Catalog of Fishes ( gen. , sp. ) | ITIS | CoL | WoRMS | Cloffa

Elasmobranchii (sharks and rays) > Rajiformes (Skates and rays) > Rajidae (Skates) > Rajinae
Etymology: Raja: Latin, raja, -ae = a sting ray (Raja sp.) (Ref. 45335 ) .  More on author: Linnaeus .

Environment: milieu / climate zone / depth range / distribution range Ecology

Marine; demersal; depth range 5 - 1020 m (Ref. 114953 ).   Subtropical; 70°N - 29°S, 25°W - 42°E

Distribution Countries | FAO areas | Ecosystems | Occurrences | Point map | Introductions | Faunafri

Eastern Atlantic and Southwest Indian Ocean: Iceland to Madagascar, including the Mediterranean Sea.

Length at first maturity / Size / Weight / Age

Maturity: L m 76.6 , range 47 - 87.5 cm
Max length : 105 cm TL male/unsexed; (Ref. 53748 ); 139.0 cm TL (female); common length : 85.0 cm TL male/unsexed; (Ref. 9987 ); max. published weight: 18.0 kg (Ref. 35388 ); max. reported age: 15 years (Ref. 41305 )

Short description Morphology | Morphometrics

Dorsal spines (total): 0; Anal spines : 0; Anal soft rays : 0. Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246 ). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167 ). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167 , 58137 ).

Biology     Glossary (e.g. epibenthic)

Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426 , 82319 , 88187 ), and from 300-577 m in the eastern Ionian Sea (Ref. 56504 ); most common in coastal waters between 10-60 m depth (Ref. 58137 ). Tolerates low salinities (Ref. 88171 ). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808 , 58137 ). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167 ) and fishes (Ref. 114953 ). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017 , 89018 ), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (<10 m) in spring (Ref. 82319 ). Young are non-migratory, inhabiting inshore nursery grounds (Ref. 89019 , 89020 , 89022 ); in the Bay of Douarnenez (France) they are found to remain in shallow waters for at least 2 years (Ref. 89023 ). Maturity age 7-8 years (Ref. 114953 ). Oviparous. Young may tend to follow large objects, such as their mother (Ref. 205 ). Detects weak electric fields generated by other organisms (prey detection and predator avoidance) but may also generate its own weak electric fields (Ref. 10311 , 88171 ). Utilized fresh and frozen (Ref. 9987 ).

Life cycle and mating behavior Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae

Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024 ). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205 , 89025 ). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167 ), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603 , 31302 ). In northwestern Europe, egg cases are laid during spring (Ref. 3167 ) and in the Mediterranean during winter and spring (Ref. 3167 ). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205 ). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250 , 88187 ) and 3.4-6.8 cm wide (Ref. 41250 ). Egg cases are anchored with an adhesive film (Ref. 82399 ). Embryos feed solely on yolk (Ref. 50449 ). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864 ). Mating season from February to September, peaking in June (Ref. 74501 ). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603 , 58137, 74501 ). Mating does not occur in the Baltic Sea (Ref. 82311 ).

Main reference Upload your references | References | Coordinator : McEachran, John | Collaborators

McEachran, J.D. and K.A. Dunn , 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314 )

IUCN Red List Status (Ref. 120744 )

  Near Threatened (NT)  ; Date assessed: 03 October 2005

CITES (Ref. 115941 )

Not Evaluated

CMS (Ref. 116361 )

Not Evaluated

Threat to humans

  Harmless




Human uses

Fisheries: commercial; gamefish: yes FAO(fisheries: production ; publication : search ) | FishSource | Sea Around Us

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Internet sources

Aquatic Commons | BHL | Cloffa | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes ( gen. , sp. ) | DiscoverLife | DORIS | ECOTOX | Faunafri | Fishtrace | GenBank( genome , nucleotide ) | GloBI | Google Books | Google Scholar | Google | IGFA World Record | MitoFish | National databases | Otolith Atlas of Taiwan Fishes | Public aquariums | PubMed | Reef Life Survey | Tree of Life | Wikipedia( Go , Search ) | World Records Freshwater Fishing | Zoobank | Zoological Record

Estimates of some properties based on models

Preferred temperature (Ref. 115969 ): 7.1 - 15.8, mean 10.7 (based on 474 cells). Phylogenetic diversity index (Ref. 82805 ):  PD 50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high]. Bayesian length-weight: a=0.00240 (0.00201 - 0.00286), b=3.24 (3.20 - 3.28), in cm Total Length, based on LWR estimates for this species (Ref. 93245 ). Trophic Level (Ref. 69278 ):  3.8   ±0.2 se; Based on diet studies. Resilience (Ref. 69278 ):  Low, minimum population doubling time 4.5 - 14 years (K=0.09-0.14; tm=10; tmax=23; Fec=150). Prior r = 0.18, 95% CL = 0.07 - 0.45, Based on 4 stock assessments. Vulnerability (Ref. 59153 ):  High to very high vulnerability (73 of 100) . Price category (Ref. 80766 ):   Medium .

Entered by Luna, Susan M. Modified by Capuli, Estelita Emily Fish Forum
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