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Gymnocarpium dryopteris (Linnaeus) Newman
WESTERN OAKFERN
Life   Plantae   Pteridophyta   Dryopteridaceae   Gymnocarpium

Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

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Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris
Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris
Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris
Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris
Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris
Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris
Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris
Gymnocarpium dryopteris
© Copyright Malcolm Storey 2011-2118 · 3
Gymnocarpium dryopteris

Associates · map
FamilyScientific name @ source (records)
Pucciniastraceae  Hyalopsora aspidiotus @ BPI (8)

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Following modified from Plants Database, United States Department of Agriculture
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http://plants.usda.gov/java/profile?symbol=GYDR ---> https://plants.usda.gov/java/profile?symbol=GYDR
https://plants.usda.gov/java/profile?symbol=GYDR ---> http://plants.usda.gov/core/profile?symbol=GYDR
http://plants.usda.gov/core/profile?symbol=GYDR ---> https://plants.usda.gov/core/profile?symbol=GYDR
&pull 20q v5.145 20180528: Error 500 read timeout https://plants.usda.gov/core/profile?symbol=GYDR

Following modified from Flora of North America
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FNA | Family List | FNA Vol. 2 | Dryopteridaceae | Gymnocarpium

3. Gymnocarpium dryopteris (Linnaeus) Newman, Phytologist. 4: app. 24. 1851.

Common oak fern, gymnocarpe fougère-du-chêne

Polypodium dryopteris Linnaeus, Sp. Pl. 2: 1093. 1753; Dryopteris linnaeana C. Christensen; Lastrea dryopteris (Linnaeus) Bory; Phegopteris dryopteris (Linnaeus) Fée; Thelypteris dryopteris (Linnaeus) Slosson

Stems 0.5--1.5 mm diam.; scales 1--4 mm. Fertile leaves usually 12--42 cm. Petiole 9--28 cm, with sparse glandular hairs distally; scales 2--6 mm. Blade broadly deltate, 2-pinnate-pinnatifid, 3--14 cm, lax and delicate, abaxial surface and rachis glabrous or with sparse glandular hairs, adaxial surface glabrous. Pinna apex entire, rounded. Proximal pinnae 2--12 cm, ± perpendicular to rachis, with basiscopic pinnules ± perpendicular to costa; basal basiscopic pinnule usually sessile, pinnatifid or rarely pinnate-pinnatifid, if sessile then with basal basiscopic pinnulet often equaling or longer than adjacent pinnulet; 2d basal basiscopic pinnule sessile, with basal basiscopic pinnulet equaling or longer than adjacent pinnulet; basal acroscopic pinnule sessile, with basal basiscopic pinnulet longer than or equaling adjacent pinnulet. Pinnae of 2d pair usually sessile with basal basiscopic pinnule longer than or equaling adjacent pinnule and about equal to basal acroscopic pinnule; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule, often with entire, rounded apex. Pinnae of 3d pair sessile with basal basiscopic pinnule equaling adjacent pinnule and equaling basal acroscopic pinnules; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule. Ultimate segments of proximal pinnae oblong, entire to crenate, apex entire, rounded. Spores 34--39 µm. 2 n = 160.

Cool, coniferous and mixed woods and at base of shale talus slopes; 0--3000 m; Greenland; St. Pierre and Miquelon; Alta., B.C., Man., N.B., Nfld., N.W.T., N.S., Ont., P.E.I., Que., Sask., Yukon; Alaska, Ariz., Colo., Conn., Idaho, Iowa, Maine, Mass., Mich., Minn., Mont., N.H., N.J., N.Mex., N.Y., Ohio, Oreg., Pa., R.I., S.Dak., Vt., Wash., W.Va., Wis., Wyo.; n,c Europe; n Asia to China, Japan.

Gymnocarpium dryopteris is a fertile allotetraploid species that arose following hybridization between G . appalachianum and G . disjunctum (see reticulogram). Its wide distribution over much of the north temperate zone has provided ample opportunity for secondary contact between G . dryopteris and each of its diploid parents, thereby resulting in a wide-ranging composite of abortive-spored triploid crosses ( G . disjunctum × G . dryopteris and G . appalachianum × G . dryopteris ). These relationships are shown on the diagram. Sterile triploid plants are not restricted only to areas where the range of the tetraploid overlaps with that of either diploid. Their broad distribution could be explained in part by their spores, which are of two types: malformed, black, and with very exaggerated perispores, or round with extensive netted perispores (K. M. Pryer and D. M. Britton 1983). The latter spore type is capable of germination and presumably permits the plants to reproduce apogamously. The name G . × brittonianum (Sarvela) Pryer & Haufler has been applied to the G . disjunctum × G . dryopteris hybrid formula (K. M. Pryer and C. H. Haufler 1993). The type of G . × brittonianum has aborted and round spores, and leaves that strongly resemble those of G . disjunctum . They are large, 3-pinnate-pinnatifid, and the second and third pairs of pinnae are sessile with basal basiscopic pinnules markedly longer than the basal acroscopic pinnules. Sterile triploid plants with a morphology similar to the type of G . × brittonianum are frequent. The biology of both of these cryptic hybrid taxa needs further study, which should lead to detailed morphologic descriptions and distribution maps.

Gymnocarpium dryopteris also hybridizes with both G . jessoense subsp. parvulum and G . robertianum .

Following modified from CalPhotos
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http://calphotos.berkeley.edu/cgi/img_query?query_src=dl&where-taxon=Gymnocarpium+dryopteris&where-lifeform=specimen_tag&rel-lifeform=ne&rel-taxon=begins+with&where-lifeform=Plant ---> https://calphotos.berkeley.edu/cgi/img_query?query_src=dl&where-taxon=Gymnocarpium+dryopteris&where-lifeform=specimen_tag&rel-lifeform=ne&rel-taxon=begins+with&where-lifeform=Plant

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Number of matches : 24
Query: SELECT * FROM img WHERE ready=1 and taxon like "Gymnocarpium dryopteris%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

Click on the thumbnail to see an enlargement

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0306 1674 [detail]
© 2006 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0306 1675 [detail]
© 2006 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0306 1676 [detail]
© 2006 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0107 0873 [detail]
© 2007 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0107 0874 [detail]
© 2007 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0107 0875 [detail]
© 2007 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0107 0876 [detail]
© 2007 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0107 0877 [detail]
© 2007 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 0000 0616 1795 [detail]
© 2016 Adrienne Simmons

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 3333 1111 1107 0058 [detail]
© University and Jepson Herbaria

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 3333 1111 1107 0059 [detail]
© University and Jepson Herbaria

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 3333 1111 1107 0071 [detail]
© University and Jepson Herbaria

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 3333 1111 1107 0072 [detail]
© University and Jepson Herbaria

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0510 2481 [detail]
© 2010 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
ID: 0000 0000 0510 2482 [detail]
© 2010 Louis-M. Landry

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3840 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3841 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3842 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3843 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3844 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3845 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3846 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3847 [detail]
© 2016 Keir Morse

Gymnocarpium dryopteris
Gymnocarpium dryopteris
Western Oak Fern
ID: 0000 1111 1234 3848 [detail]
© 2016 Keir Morse

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


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