Hedwig, Sp. Musc. Frond. 155, plate 39, figs. 1-5. 1801.
to 11 × 3.5 mm.
mostly branched; axillary hyaline nodules absent; central strand present.
as many as 15 pairs, often undulate, oblong to lanceolate, obtuse to broadly acute, cuspidate, to 2.7 mm × 0.6 mm; dorsal lamina truncate-rounded proximally, ending at insertion, not decurrent; vaginant laminae
/3 the leaf length, ± equal, minor lamina ending on or near margin; margin evenly serrulate, often crenulate-serrulate on dorsal and ventral laminae, elimbate; costa ending in cuspidate apex, taxifolius-type; laminal cells 1-stratose, distinct, smooth, bulging, firm-walled, irregularly hexagonal, 8-11 µm frequently paler at margin, conspicuously mammillose in vaginant laminae, larger and clearer juxtacostally.
rhizautoicous; perigonia on short branches proximal to elongate stems; perichaetia on short, proximal, axillary branches.
1 per perichaetium.
to 17 mm.
theca slightly inclined, slightly arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum to 1 mm.
cucullate, smooth, to 2 mm.
Damp, shaded soil, humus, and rocks; N.W.T., N.S., Ont., P.E.I., Que.; Ala., Ark., Ariz., Calif., Del., D.C., Fla., Ga., Ill., Ind., Iowa, Kans., Ky., La., Maine, Mass., Mich., Miss., Mo., Nebr., N.H., N.J., N.Y., N.C., Ohio, Okla., Oreg., Pa., S.C., Tenn., Tex., Vt., Va., Wash., W.Va., Wis.; Mexico; West Indies; Central America; South America (Brazil and Chile); Europe; Asia (China, Japan); Pacific Islands ( New Zealand).
is distinguished by evenly serrulate or crenulate-serrulate leaf margin, stout costa ending in leaf cusp, conspicuous mammillose cells in the vaginant laminae, and sporophytes on short branches in the axils of proximal leaves. It can be confused with
, but that species is smaller, has costa shorter, and cells in the vaginant laminae have small, more or less inconspicuous papillae in the corners of the walls.
usually occurs on shaded, damp soil or humus, whereas
is usually found on disturbed soil in woods, along paths, and in road cuts.
is considered to be a freakish expression of
(R. A. Pursell 2003).