|California Fungi—Dendrocollybia racemosa
(Photo: © Michael Wood)
(Pers.) R.H. Petersen & S. A. Redhead
Mycol. Res. 105(2): 169.
Cap 0.7-1.5 cm in diameter, broadly conic, becoming convex, eventually plano-convex, the disc slightly umbonate; (in some specimens the cap may be greatly reduced or absent); margin decurved, becoming plane to raised, entire, eroded or wavy; surface dull brown, glabrous at the disc, elsewhere appressed silky-fibrillose, grey-brown, paler towards the margin, sometimes faintly zonate; context thin, less than 1.0 mm thick, grey; odor, not distinctive; taste mild.
Gills adnexed to notched, close, moderately broad, dingy greyish-tan; lamellulae up to 3-seried.
Stipe 4.0-6.0 cm long, 0.5-1.0 mm thick, more or less equal, pliant, hollow to stuffed; surface of apex dull tan-brown, pruinose, darker brown below, covered with scattered lighter colored fibrils, short side branches with swollen tips projecting from the lower two-thirds of the stipe, the stipe often well buried in the substrate.
Sexual spores 4.0-4.5 x 2.0-2.5 µm, ellipsoid, thin-walled, hilar appendage conspicuous, non-amyloid; spore print white. Asexual spores 10.0-15.5 x 3-4 µm, narrowly ellipsoid to oblong, contents granular, non-amyloid.
Solitary or in small groups growing from a grain-like sclerotium on the decayed remains of decayed mushrooms, or in duff of mixed hardwood-conifer woods; fruiting from late fall to mid-winter.
This easily overlooked small, greyish mushroom is recognized by a stipe with many short, side branches. The branches which end in swollen tips produce asexual spores, presumably a hedge to the sexual spores of the cap. In some specimens asexual reproduction predominates with the cap either greatly reduced or absent. A cousin,
, is also small and inconspicuous. Like
, it grows from a sclerotium on the remains of decayed mushrooms, but can be distinguished by a whitish cap, and the lack of lateral stipe branches.
Bas, C., Kyper, T.W., Noordeloos, M.E. & Vellinga, E.C.
(1995). Flora Agaricina Neerlandica -- Critical monographs on the families of agarics and boleti occuring in the Netherlands. Volume 3. Tricholomataceae. A. A. Balkema: Rotterdam, Netherlands. 183 p.
Castellano, M.A., Cázares, E., Fondrick, B. & Dreisbach, T.
(2003). Handbook to additional fungal species of special concern in the Northwest Forest Plan (Gen. Tech Rep. PNW-GTR-572). U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station: Portland, OR. 144 p. (
Desjardin, D.E., Wood, M.G. & Stevens, F.A.
(2015). California Mushrooms: The Comprehensive Identification Guide. Timber Press: Portland, OR. 560 p.
Hughes, K.W., Petersen, R.H., Johnson, J.E., Moncalvo, J.-M. , Vilgalys, R., Redhead, S.A., Thomas, T. & McGhee, L.A.
(2001). Infragenic phylogeny of
s. str. based on sequences of ribosomal ITS and LSU regions. Mycol. Res. 105(2): 164-172.
(1979). Collybioid genera in the Pacific Northwest. Mycotaxon 9(1): 117-231.
Siegel, N. & Schwarz, C.
(2016). Mushrooms of the Redwood Coast. Ten Speed Press: Berkeley, CA. 601 p.
Siegel, N., Vellinga, E.C., Schwarz, C., Castellano, M.A. & Ikeda, D.
(2019). A Field Guide to the Rare Fungi of California's National Forests. Bookmobile: Minneapolis, MN. 313 p. (
Other Descriptions and Photos
(D & CP)
Jens H. Petersen:
Arora (1986): p. 213 (D), p. 212 (P) [as
Smith (1975): sp. 109 (D & CP) [as
(D=Description; I=Illustration; P=Photo; CP=Color Photo)
The Fungi of California
Copyright © 2000-2019 Michael Wood & Fred Stevens